Insectivorous birds
- Tits especially pick overwintering larvae from bark crevices, but do not forage specifically for the pest unless population densities are very high and for this reason are of limited value only.
Egg parasites
- The egg parasitic wasp Trichogramma can be introduced (4 releases of 2.5 m per ha have been shown to reduce damage by 40-85%) but such introductions are not cost effective.
Larval parasites
- The parasitic wasp Colpoclypeus florus is an external parasite of third to fifth instar summer fruit tortrix moth larvae.
- The parasite can be seen attached behind the head of its host.
Predatory insects
- Earwigs and predatory mirid and anthocorid bugs feed of eggs and young larvae.
Virus diseases
- A nucleopolyhedrovirus (AoNPV) and two strains of a granulovirus (AoGV) of summer fruit tortrix moth are known (see ‘Biological control’).
- These are normally found in association with commercial applications of biocontrol agents, though natural infections of AoGV were found in two orchards near Faversham in Kent in 1993.