The commonest species on apple, Edwardsiana crataegi, overwinters as eggs laid in the autumn beneath the bark of twigs or small branches of fruit trees.
- Eggs hatch in April to early May.
- Nymphs feed until mid-summer.
- New adults appear in late July and August, laying eggs in or near the mid-vein on the undersides of leaves.
- Second generation nymphs occur from August onwards and become adults in the autumn.
Edwardsiana rosae has a similar life cycle but overwintering eggs are laid on rose.
- Apple orchards are invaded by a migration of adults to summer hosts in June.
- There is a return migration of second generation adults to rose to lay eggs in the autumn.